Scottish daily The Scotsman and its sister Sunday title reach more than one million people each month in print and online, according to the latest readership figures.
The National Readership Survey has published new figures combining the estimated print and website audiences for national newspapers and a handful of regional titles.
It shows that The Scotsman and Scotland on Sunday had combined print and online readers of 1.1m and around half of these came from its website, which was used by 582,000 people.
The NRS Print and Digital Data figures showed there were 447,000 print readers of The Scotsman a month and when combined with the print readers of Scotland on Sunday, this increased to 592,000.
The NRS figures, which are estimated from surveys with individuals rather than sales figures, relate to the whole of 2012 and the online data is provided by UKOM/comScore for November 2012.
Of the other titles in the survey, Glasgow-based The Herald and the Sunday Herald reached an estimated 836,000 people a month between them, 359,000 of which accessed its website.
The Yorkshire Post had a combined print and online audience of 792,000 and 166,000 of these were website users, while Dundee-based The Courier had a combined monthly readership of 326,000, with 55,000 of these being digital users.
The online figures are calculated for the first time using data from comScore, meaning it is not possible to compare the latest combined readership figures with previous ones released last year.
These titles are currently the only regional ones covered by the NRS in its 36,000 face-to-face interviews carried out each year but from next month it will start collecting data for all regional newspapers in Britain.