Mr Pelosi is to remain with the company until the end of the year, but in a different role working with A&N Media chief executive Kevin Beatty on group-wide initiatives.
Also standing down is Mr Pelosi’s deputy, Alex Leys, who is retiring after a 28-year career with the company which included several regional editorships.
Linda Grant, currently group commercial development director, will replace Mr Auckland at Metro.
All are due to take up their new posts after a short handover period.
Announcing the move to staff, Mr Beatty said: “There will be opportunities in the future to acknowledge the huge contribution that both Michael and Alex have made to the group.
“In the meantime I am sure you will join me in wishing them all every success.”